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a    nR_eSNV
w    Harpalus discrepans Morawitz, 1862

̏Wn kCΎs 1993/7/18

      z kC, {B, l, B
ԏ _kn ({c 1980), n (}ER 1990), ͐~ (Ishitani et al. 1997), X (Siddiquee and Nakamura 2004, Suttiprapan and Nakamura 2007), _n (Salah et al. 2004), Lt ({ 2005, { 2008, { 2009a), ss (Fujita et al. 2008), Gؗ ({ 2009a), ͐~ (EΈ 2010), Ɨt ({ 2009b), (Habu 1973d), ł4{5{ (Habu 1973d)
^Cv @ @ @
      l Harpalus modestus niponensis Bates (Habu 1968c), Harpalus discrepans Morawitz, Pheuginus corporosus Motschulsky, Harpalus flavitarsis Dejean var. niponensis Bates (Habu 1973d)